Medium-sized passerine, feeds on seeds, berries, and (in summer) invertebrates, especially Lepidoptera larvae and beetles (Coleoptera); in winter quarters specializes in beechnuts Fagus sylvatica. Breeds across boreal and subarctic zones of west Palearctic, in the more open birch Betula woods, and in mixed forests of birch and conifers. Winters in Europe from S Sweden to N Morocco and from UK to Turkey and Georgia. The population trend appears to be decreasing, but not sufficiently rapid to approach the thresholds for Vulnerable. A medium distance migrant. Extent of movement is strongly dependent on food availability; local concentrations of millions of birds occur in south-central Europe.